Despite their great socio-economic importance, sandy beaches have attracted little ecological research. This is unfortunate since, contrary to popular belief, they support diverse ecological assemblages whose species are mostly small and buried and which deserve protection as part of ecologically sustainable development (ESD). Moreover, the management of beaches and linked adjacent ecosystems is becoming increasingly important because of their vulnerability to burgeoning human pressures including climate change. Although there are large uncertainties involved, some of the climate-related environmental changes and their ecological consequences for sandy beaches are explored in this paper, some management issues discussed and research proposed.

Temperature-related changes include the likelihood that the geographical ranges of some species will change, some cool-adapted species will decline in abundance, possibly to extinction, and the rate of processes such as decomposition and photosynthesis will increase. The increasing acidification of the ocean may affect many beach species directly via reduced calcification and indirectly via changes to the phytoplankton on which some beach species depend for food. Changed hydrology may affect both nearshore primary productivity and the larval movements of many species. Increased storminess may change the morphodynamic state of beaches with consequent effects on the diversity, abundance and composition of beach assemblages. The combination of storm surges and sea-level rise may enhance erosion rates, the most extreme effect being the total loss of sandy beach habitat, especially if sea-walls are deployed to protect coastal assets. Alternatively, in some areas there may be a slow retreat of the coastline with few effects on beach ecosystems.

Concerning management, both mitigation and adaptation strategies are needed to meet public ESD goals. The former would seek to constrain the increase in atmospheric greenhouse gases by addressing the underlying causes (i.e., population and economic growth) and by applying appropriate technologies. Adaptation strategies would recognise the linkages between beaches, dunes and surf zones and maintain sand movement and storage and allow for the landward migration of beaches. Failing this, the active maintenance of beaches in their present location by soft engineering (e.g., the transport of sand to the intertidal beach from elsewhere, a process called beach nourishment) is far preferable to hard engineering (e.g., sea-walls) from the nature conservation and socio-economic points of view. Since beach ecosystems are poorly understood even at basic descriptive levels, future management, monitoring, and predictions of the possible impacts of large-scale phenomena (such as global climate change) will be assisted by basic research into documenting biodiversity, community dynamics, autecology, and eco-physiology of beach fauna and the linkages with adjacent ecosystems. As well, the use of human interventions (e.g., beach nourishment) as scientific experiments combined with optimised monitoring is recommended.

ABS 2003. ABS Catalogue 3101.0. Australian Bureau of Statistics, Canberra.
ABS Catalogue 3101.0
Alonso, J.A-C. and Cabrera, J. 2002. Tourist resorts and their impact on beach erosion at Sotavento beaches, Fuerteventura, Spain. Journal of Coastal Research SI 36: 1-7.
Tourist resorts and their impact on beach erosion at Sotavento beaches, Fuerteventura, Spain
Journal of Coastal Research
, vol. 
36
 (pg. 
1
-
7
)
Anon 2005. Sands of time threaten beaches. The Sun Herald, 21/08/05.
Anon 2006. Deadline set on climate disaster. The Australian, 01/02/06
Ansell, A.D. 1983. The biology of the genus Donax. In Sandy Beaches as Ecosystems. Eds A. McLachlan and T. Erasmus. Pp. 607-635. Junk, The Hague.
Sandy Beaches as Ecosystems
(pg. 
607
-
635
)
Australia's Oceans Policy 1998. Integrated and Ecosystem-Based Oceans Planning and Management. www.oceans.gov.au.
Bambach, R.K., Knoll, A.H. and Sepkowski, J.J., Jr. 2002. Anatomical and ecological constraints on Phanerozoic animal diversity in the marine realm. Proceedings of the National Academy of Sciences of the United States 99: 6845-6859.
Anatomical and ecological constraints on Phanerozoic animal diversity in the marine realm
Proceedings of the National Academy of Sciences of the United States
, vol. 
99
 (pg. 
6845
-
6859
)
Barros, F. 2001. Ghost crabs as a tool for rapid assessment of human impacts on exposed sandy beaches. Biological Conservation 97: 399-404.
Ghost crabs as a tool for rapid assessment of human impacts on exposed sandy beaches
Biological Conservation
, vol. 
97
 (pg. 
399
-
404
)
Batley, G.E. and Cocks, K.D. 1992. Defining and Quantifying National Coastal Resources. Working document 92/11. CSIRO, Canberra.
Working document 92/11
Berner, R.A. 2002. Examination of hypotheses for the Permo-Triassic boundary extinction by carbon cycle modeling. Proceedings of the National Academy of Sciences of the United States 99: 4172-4177.
Examination of hypotheses for the Permo-Triassic boundary extinction by carbon cycle modeling
Proceedings of the National Academy of Sciences of the United States
, vol. 
99
 (pg. 
4172
-
4177
)
Bilodeau, A.L. and Bourgeois, R.P. 2004. Impact of beach restoration on the deep-burrowing Ghost Shrimp, Callichirus islagrande. Journal of Coastal Research 20: 931-936.
Impact of beach restoration on the deep-burrowing Ghost Shrimp, Callichirus islagrande
Journal of Coastal Research
, vol. 
20
 (pg. 
931
-
936
)
Bird, E.C.F. 1996. Beach Management Chichester, UK: Wiley.
Beach Management
Braziero, A. and Defeo, O. 1999. Effects of harvesting and density-dependence on the demography of sandy beach populations: the yellow clam Mesodesma mactroides of Uruguay. Marine Ecology Progress Series 182: 127-135.
Effects of harvesting and density-dependence on the demography of sandy beach populations: the yellow clam Mesodesma mactroides of Uruguay
Marine Ecology Progress Series
, vol. 
182
 (pg. 
127
-
135
)
Blackwell, B. 1997. An environmental-economic discussion of beach erosion - its causes, effects and remedies. Queensland Conservation Group's Conference, Mackay.
Blackwell, B.D. 2003. The Economics of Coastal Foreshore and Beach Management: Use, Safe Bathing Facilities, Erosion and Conservation, PhD Thesis, School of Economics, The University of Queensland, Brisbane.
The Economics of Coastal Foreshore and Beach Management: Use, Safe Bathing Facilities, Erosion and Conservation
Blackwell, B.D. (in press). The Economic Value of Some of Australia's Natural Coastal Assets: The Value of Their Ecosystem Goods and Services: Final Report, Cooperative Research Centre for Coastal Zone, Estuary and Waterway Management, Indooroopilly, Queensland.
Bouchard, S.S. and Bjorndal, K.A. 2000. Sea turtles as biological transporters of nutrients and energy from marine to terrestrial ecosystems. Ecology 81: 2305-2313.
Sea turtles as biological transporters of nutrients and energy from marine to terrestrial ecosystems
Ecology
, vol. 
81
 (pg. 
2305
-
2313
)
Breusch, J. 2006. Greenhouse summit rejects fossil fuel cuts. Financial Review, 13/1/06, p.1.
Brazeiro, A. 2001. Relationship between species richness and morphodynamics in sandy beaches: what are the underlying factors? Marine Ecology Progress Series 224: 35-44.
Relationship between species richness and morphodynamics in sandy beaches: what are the underlying factors?
Marine Ecology Progress Series
, vol. 
224
 (pg. 
35
-
44
)
Brewer, P.G. 1997. Ocean chemistry of the fossil fuel CO2 signal: the haline signal of “business as usual”. Geophysical Research Letters 24: 1367-1369.
Ocean chemistry of the fossil fuel CO2 signal: the haline signal of “business as usual”
Geophysical Research Letters
, vol. 
24
 (pg. 
1367
-
1369
)
Brown, A.C. 2001. Biology of Sandy Beaches. In: Encyclopedia of Ocean Sciences. Eds J.H. Steele, S.A. Thorpe & K.K.Turekian. Volume 5, Pp. 2496-2504. Academic Press, London.
Encyclopedia of Ocean Sciences
(pg. 
2496
-
2504
)
Brown, A.C. and McLachlan, A. 1990. Ecology of Sandy Shores. Elsevier, Amsterdam.
Ecology of Sandy Shores
Brown, A.C. and McLachlan, A. 2002. Sandy shore ecosystems and the threats facing them: some predictions for the year 2025. Environmental Conservation 29: 62-77.
Sandy shore ecosystems and the threats facing them: some predictions for the year 2025
Environmental Conservation
, vol. 
29
 (pg. 
62
-
77
)
Brown, A.C., McLachlan, A., Kerley, G.I.H. and Lubke, R.A. 2000. Functional Ecosystems: Sandy Beaches and Dunes. In: Marine Biodiversity Status Report. Eds B.D. Durham & J.C. Pauw.
Bruun, P. 1988. The Bruun Rule of erosion by sea level rise: a discussion of large-scale two- and three- dimensional usages. Journal of Coastal Research 4: 627-648.
The Bruun Rule of erosion by sea level rise: a discussion of large-scale two- and three- dimensional usages
Journal of Coastal Research
, vol. 
4
 (pg. 
627
-
648
)
Burkett, V., Codignotto, J.O., Forbes, D.L., Mimura, N., Beamish, R.J., and Ittekkot, V. 2001. Coastal zones and marine ecosystems pp 343-380 In: Climate Change 2001: Impacts, Adaptation, and Vulnerability. Eds J.J. McCarthy, O.F. Canziani, N.A. Leary, D.J. Dokken and K.S. White. Contribution of Working Group II to the Third Assessment report of the Intergovernmental Panel on Climate Change (Cambridge University Press, Cambridge).
Climate Change 2001: Impacts, Adaptation, and Vulnerability
(pg. 
343
-
380
)
Cocks, K.D. and Crossland, C. 1991. The Australian coastal zone: a discussion paper. Working Document 91/4. CSIRO, Canberra.
The Australian coastal zone: a discussion paper
Colombini, I. and Chelazzi, L. 2003. Influence of marine allochthonous input on sandy beach communities. Oceanography and Marine Biology Annual Review 41: 115-159.
Influence of marine allochthonous input on sandy beach communities
Oceanography and Marine Biology Annual Review
, vol. 
41
 (pg. 
115
-
159
)
Council of Australian Governments 1992. National Strategy for Ecologically Sustainable Development. AGPS, Canberra.
National Strategy for Ecologically Sustainable Development
Cowell, P.J. and Thom, B.G. 1994. Coastal impacts of climate change - modelling procedures for use in local government. Proc. 1st National Coastal Management Conference, Coast to Coast '94, Hobart. Pp. 43-50.
Cowell, P.J., Thom, B.J., Jones, R.A., Everts, C.H. and Simanovic, D. 2006. Management of uncertainty in predicting climate-change impacts on beaches. Journal of Coastal Research 22: 232-245.
Management of uncertainty in predicting climate-change impacts on beaches
Journal of Coastal Research
, vol. 
22
 (pg. 
232
-
245
)
Defeo, O. 1998. Testing hypotheses on recruitment, growth and mortality in exploited bivalves: an experimental perspective. Canadian Special Publications on Fisheries and Aquatic Sciences 125: 257-264.
Testing hypotheses on recruitment, growth and mortality in exploited bivalves: an experimental perspective
Canadian Special Publications on Fisheries and Aquatic Sciences
, vol. 
125
 (pg. 
257
-
264
)
Defeo, O., Lercari, D. and Gomez, J. 2003. The role of morphodynamics in structuring sandy beach populations and communities: What should be expected? Journal of Coastal Research 35: 352-362.
The role of morphodynamics in structuring sandy beach populations and communities: What should be expected?
Journal of Coastal Research
, vol. 
35
 (pg. 
352
-
362
)
Denholm, M. 2006. Studies back fears sea levels are rising rapidly. The Australian, 1/2/06
Dexter, D.M. 1983a. Community structure of intertidal sandy beaches in New South Wales, Australia. In: Sandy Beaches as Ecosystems. Eds A. McLachlan and T. Erasmus. Pp. 461-472. Junk, The Hague.
Sandy Beaches as Ecosystems
(pg. 
461
-
472
)
Dexter, D.M. 1983b. A guide to the sandy beach fauna of New South Wales. Wetlands 3: 94-104.
A guide to the sandy beach fauna of New South Wales
Wetlands
, vol. 
3
 (pg. 
94
-
104
)
Dexter, D.M. 1984. Temporal and spatial variability in the community structure of the fauna of four sandy beaches of south-eastern New South Wales. Australian Journal of Marine and Freshwater Research 35: 663-672.
Temporal and spatial variability in the community structure of the fauna of four sandy beaches of south-eastern New South Wales
Australian Journal of Marine and Freshwater Research
, vol. 
35
 (pg. 
663
-
672
)
Dexter, D.M. 1985. Distribution and life histories of abundant crustaceans of four sandy beaches on south-eastern New South Wales. Australian Journal of Marine and Freshwater Research 36: 281-289.
Distribution and life histories of abundant crustaceans of four sandy beaches on south-eastern New South Wales
Australian Journal of Marine and Freshwater Research
, vol. 
36
 (pg. 
281
-
289
)
DLWC 2001. Coastal Dune Management: A Manual of Coastal Dune Management and Rehabilitation Techniques. NSW Department of Land and Water Conservation, Coastal Unit, Newcastle, Australia.
Coastal Dune Management: A Manual of Coastal Dune Management and Rehabilitation Techniques
Donn, T.E. 1987. Longshore distribution of Donax serra in two log-spiral bays in eastern Cape, South Africa. Marine Ecology Progress Series 35: 217-222.
Longshore distribution of Donax serra in two log-spiral bays in eastern Cape, South Africa
Marine Ecology Progress Series
, vol. 
35
 (pg. 
217
-
222
)
Dovers, S.R. and Handmer, J.W. 1995. Ignorance, the precautionary principle and sustainability. Ambio 24: 92-97.
Ignorance, the precautionary principle and sustainability
Ambio
, vol. 
24
 (pg. 
92
-
97
)
Dugan, J.E., Hubbard, D.M., McCrary, M.D. and Pierson, M.O. 2003. The response of macrofauna communities and shorebirds to macrophyte wrack subsidies on exposed sandy beaches of southern California. Estuarine, Coastal and Shelf Science 58 S: 25-40.
The response of macrofauna communities and shorebirds to macrophyte wrack subsidies on exposed sandy beaches of southern California
Estuarine, Coastal and Shelf Science
, vol. 
58 S
 (pg. 
25
-
40
)
Dugan, J.E., Jaramillo, E., Hubbard, D.M., Contreras, H. and Duarte, C. 2004. Competitive interactions in macroinfaunal animals of exposed sandy beaches. Oecologia 139: 630-640.
Competitive interactions in macroinfaunal animals of exposed sandy beaches
Oecologia
, vol. 
139
 (pg. 
630
-
640
)
Du Preez, H.H., McLachlan, A., Marais, J.F.K. and Cockroft, A.C. 1990. Bioenergetics of fishes in a high-energy surf-zone. Marine Biology 106: 1-12.
Bioenergetics of fishes in a high-energy surf-zone
Marine Biology
, vol. 
106
 (pg. 
1
-
12
)
Edgar, G.J., Moverly, J., Barrett, N.S., Peters, D. and Reed, C. 1997. The conservation-related benefits of a systematic sampling programme: the Tasmanian reef bioregionalisation as a case study. Biological Conservation 79: 227-240.
The conservation-related benefits of a systematic sampling programme: the Tasmanian reef bioregionalisation as a case study
Biological Conservation
, vol. 
79
 (pg. 
227
-
240
)
Fairweather, P.G. 1990. Ecological changes due to our use of the coast: research needs versus effort. Proc. Ecological Society Australia 16: 71-77.
Ecological changes due to our use of the coast: research needs versus effort
Proc. Ecological Society Australia
, vol. 
16
 (pg. 
71
-
77
)
Feely, R.A., Sabine, C.L., Lee, K., Berelson, W., Kleypas, J., Fabry, V.J., and Millero, F.J. 2004. Impact of anthropogenic CO2 on the CaCO3 system in the oceans. Science 305: 362-366.
Impact of anthropogenic CO2 on the CaCO3 system in the oceans
Science
, vol. 
305
 (pg. 
362
-
366
)
Finkl, C.W. and H.J. Walker (2004). Beach Nourishment. In: Schwartz, M. (ed.). The Encyclopedia of Coastal Science, Dordrecht, The Netherlands, Kluwer Academic Press (noe Springer). Pp. 37-54.
The Encyclopedia of Coastal Science
(pg. 
37
-
54
)
Gladstone W., Hacking N., Owen V. (2006). Effects of artificial openings of intermittently opening estuaries on assemblages of macroinvertebrates in the entrance barrier. Estuarine, Coastal and Shelf Science 67: 708-720.
Effects of artificial openings of intermittently opening estuaries on assemblages of macroinvertebrates in the entrance barrier
Estuarine, Coastal and Shelf Science
, vol. 
67
 (pg. 
708
-
720
)
Gray, J.S. 1974. Animal-sediment relationships. Oceanography and Marine Biology Annual Review 12: 223-261.
Animal-sediment relationships
Oceanography and Marine Biology Annual Review
, vol. 
12
 (pg. 
223
-
261
)
Griffiths, C.L. and Stenton-Dozey, J. 1981. The fauna and rate of degradation of stranded kelp. Estuarine, Coastal and Shelf Science 12: 645-653.
The fauna and rate of degradation of stranded kelp
Estuarine, Coastal and Shelf Science
, vol. 
12
 (pg. 
645
-
653
)
Grumbine, R.E. 1994. What is ecosystem management? Conservation Biology 8: 27-38.
What is ecosystem management?
Conservation Biology
, vol. 
8
 (pg. 
27
-
38
)
Hacking, N. 1996. Tidal movement of sandy beach macrofauna. Wetlands (Australia) 15: 55-71.
Tidal movement of sandy beach macrofauna
Wetlands (Australia)
, vol. 
15
 (pg. 
55
-
71
)
Hacking, N. 1998. Macrofaunal community structure of beaches in northern New South Wales, Australia. Marine and Freshwater Research 49: 47-53.
Macrofaunal community structure of beaches in northern New South Wales, Australia
Marine and Freshwater Research
, vol. 
49
 (pg. 
47
-
53
)
Harley, C.D.G., Hughes, A.R., Hultgren, K.M., Miner, B.G., Sorte, C.J.B., Thornber, C.S., Rodriguez, L.F., Tomanek, L and Williams, S.L. 2006. The impacts of climate change in global marine systems. Ecology Letters 9: 228-241.
The impacts of climate change in global marine systems
Ecology Letters
, vol. 
9
 (pg. 
228
-
241
)
Harper, E.M. 2000. Are calcitic layers an effective adaptation against shell dissolution in the Bivalvia? Journal of Zoology London 251: 179-186.
Are calcitic layers an effective adaptation against shell dissolution in the Bivalvia?
Journal of Zoology London
, vol. 
251
 (pg. 
179
-
186
)
Haugan, P.M. and Drange, H. 1996. Effects of CO2 on the ocean environment. Energy Conversion Management 37: 1019-1022.
Effects of CO2 on the ocean environment
Energy Conversion Management
, vol. 
37
 (pg. 
1019
-
1022
)
Haynes, D. and Devlin, M. 1997. The sandy intertidal: a neglected marine eco system. In: Aust.Coral Reef Soc.75th Anniv. Conf. J.G. Greenwood and N. Hall (eds).
Healey, M.C. and Hennessey, T.M. 1994. The utilization of scientific information in the management of estuarine information. Ocean and Coastal Management 23:167-191.
The utilization of scientific information in the management of estuarine information
Ocean and Coastal Management
, vol. 
23
 (pg. 
167
-
191
)
Hughes, L. 2003. Climate change and Australia: trends, projections and impacts. Austral Ecology 28: 423-443.
Climate change and Australia: trends, projections and impacts
Austral Ecology
, vol. 
28
 (pg. 
423
-
443
)
International Scientific Steering Committee 2005. Avoiding Dangerous Climate Change. Report presented at: International Symposium on the Stabilisation of Greenhouse Gas Concentrations. Exeter, UK. 1-3 February 2005.
IPCC 2001. Intergovernmental Panel on Climate Change web site. www.grida.no/climate/ipcc_tar/wg2/index.htm
James, R.J. 2000a. From beaches to beach environments: linking the ecology, human-use and management of beaches in Australia. Ocean & Coastal Management43: 495-514.
From beaches to beach environments: linking the ecology, human-use and management of beaches in Australia
Ocean & Coastal Management
, vol. 
43
 (pg. 
495
-
514
)
James, R.J. 2000b. The first step for the environmental management of Australian beaches: establishing an effective policy framework. Coastal Management 28: 149-160.
The first step for the environmental management of Australian beaches: establishing an effective policy framework
Coastal Management
, vol. 
28
 (pg. 
149
-
160
)
James, R.J. and Fairweather, P.G. 1996. Spatial variation of intertidal macrofauna on a sandy ocean beach in Australia. Estuarine Coastal and Shelf Science 43: 81-107.
Spatial variation of intertidal macrofauna on a sandy ocean beach in Australia
Estuarine Coastal and Shelf Science
, vol. 
43
 (pg. 
81
-
107
)
Janzen, F.J. 1994. Climate-change and temperature -dependent sex determination in reptiles. Proceedings of the National Academy of Sciences, USA. 91: 7484-7490.
Climate-change and temperature -dependent sex determination in reptiles
Proceedings of the National Academy of Sciences, USA
, vol. 
91
 (pg. 
7484
-
7490
)
Jaramillo, E. and McLachlan, A. 1993. Community and population responses of the macroinfauna to physical factors over a range of exposed sandy beaches in South-central Chile. Estuarine Coastal and Shelf Science 37: 615-624.
Community and population responses of the macroinfauna to physical factors over a range of exposed sandy beaches in South-central Chile
Estuarine Coastal and Shelf Science
, vol. 
37
 (pg. 
615
-
624
)
Jaramillo, E., Contreras, H. and Bollinger, A. 2002. Beach and faunal response to the construction of a seawall in a sandy beach of south central Chile. Journal of Coastal Research 18: 523-529
Beach and faunal response to the construction of a seawall in a sandy beach of south central Chile
Journal of Coastal Research
, vol. 
18
 (pg. 
523
-
529
)
Jones, A.R. and Short, A.D. 1995. Sandy Beaches. Chap. 9 in Coastal Marine Ecology of Temperate Australia. Eds A.J. Underwood and M.G. Chapman. University of New South Wales Press, Sydney. 341pp.
Coastal Marine Ecology of Temperate Australia
Jones, A.R., Murray, A. and Marsh, R.E. 1991. Patterns of abundance of Exoedicerotid amphipods on sandy beaches near Sydney, Australia. Hydrobiologia 223: 119-126.
Patterns of abundance of Exoedicerotid amphipods on sandy beaches near Sydney, Australia
Hydrobiologia
, vol. 
223
 (pg. 
119
-
126
)
Jones, A.R., Murray, A. and Marsh, R.E. 1998. A method for sampling sandy beach amphipods that tidally migrate. Marine and Freshwater Research 49: 863-865.
A method for sampling sandy beach amphipods that tidally migrate
Marine and Freshwater Research
, vol. 
49
 (pg. 
863
-
865
)
Kennedy, V.S., Twilley, R.R., Kleypas, J.A., Cowan, J.H., Jr. and Hare, S.R. 2002. Coastal and Marine Ecosystems & Global Climate Change: Potential Effects on U.S. Resources. Prepared for the Pew Center on Global Climate Change, Arlington, VA.
Coastal and Marine Ecosystems & Global Climate Change: Potential Effects on U.S. Resources
Kennett, J.P. and Stott, L.D. 1991. Abrupt deep-sea warming, palaeoceanographic changes and benthic extinctions at the end of the Palaeocene. Nature 353: 225-229.
Abrupt deep-sea warming, palaeoceanographic changes and benthic extinctions at the end of the Palaeocene
Nature
, vol. 
353
 (pg. 
225
-
229
)
Koop, K., Newell, R.C. and Lucas, M.I. 1982. Biodegradation and carbon flow based on kelp (Ecklonia maxima) debris in a sandy beach microcosm. Marine Ecology Progress Series 7: 315-326.
Biodegradation and carbon flow based on kelp (Ecklonia maxima) debris in a sandy beach microcosm
Marine Ecology Progress Series
, vol. 
7
 (pg. 
315
-
326
)
Krebs, C.J. 1978. Ecology. The Experimental Analysis of Distribution and Abundance. Harper and Row, New York. 2nd Edition.
Ecology. The Experimental Analysis of Distribution and Abundance
Lastra, M., Jaramillo, E., López, J., Contreras, H., Duarte, C. and Rodriguez, J.G. 2004. Population abundances, tidal movement, burrowing ability and oxygen uptake of Emerita analoga (Stimpson) (Crustacea, Anomura) on a sandy beach of south-central Chile. Marine Ecology 25: 71-89.
Population abundances, tidal movement, burrowing ability and oxygen uptake of Emerita analoga (Stimpson) (Crustacea, Anomura) on a sandy beach of south-central Chile
Marine Ecology
, vol. 
25
 (pg. 
71
-
89
)
Leemans, R. and van Vliet, A. 2005. Responses of species to changes in climate determine climate protection targets. Report presented at: International Symposium on the Stabilisation of Greenhouse Gas Concentrations. Exeter, UK. 1-3 February 2005.
Lenanton, R.C.J. and Caputi, N. 1989. The role of food supply and shelter in the relationship between fishes, in particular Cnidoglanis macrocephalus Valenciennes, and detached macrophytes in the surf-zone of sandy beaches. Journal of Experimental Marine Biology and Ecology 128: 165-76.
The role of food supply and shelter in the relationship between fishes, in particular Cnidoglanis macrocephalus Valenciennes, and detached macrophytes in the surf-zone of sandy beaches
Journal of Experimental Marine Biology and Ecology
, vol. 
128
 (pg. 
165
-
76
)
Lowenstam, H.A. 1981. Minerals formed by organisms. Science 211: 1126-1131.
Minerals formed by organisms
Science
, vol. 
211
 (pg. 
1126
-
1131
)
McGwynne, L.E., McLachlan, A., and Furstenburg, J.P. 1988. Wrack break-down on sandy beaches. Its impact on interstitial meiofauna. Marine Environmental Research 25: 213-232.
Marine Environmental Research
Marine Environmental Research
, vol. 
25
 (pg. 
213
-
232
)
McLachlan, A. 1985. The biomass of macro- and interstitial fauna on clean and wrack-covered beaches in Western Australia. Estuarine Coastal and Shelf Science 21: 587-599.
The biomass of macro- and interstitial fauna on clean and wrack-covered beaches in Western Australia
Estuarine Coastal and Shelf Science
, vol. 
21
 (pg. 
587
-
599
)
McLachlan, A. 1990. Dissipative beaches and macrofauna communities on exposed intertidal sands. Journal of Coastal Research 6: 57-71.
Dissipative beaches and macrofauna communities on exposed intertidal sands
Journal of Coastal Research
, vol. 
6
 (pg. 
57
-
71
)
McLachlan, A. 1998. Interactions between two species of Donax on a high energy beach: an experimental approach. Journal of Molluscan Studies 64: 492-495.
Interactions between two species of Donax on a high energy beach: an experimental approach
Journal of Molluscan Studies
, vol. 
64
 (pg. 
492
-
495
)
McLachlan, A. and Dorvlo, A. 2004. Global patterns in sandy beach macrobenthic communities. Journal of Coastal Research 21: 674-687.
Journal of Coastal Research
Journal of Coastal Research
, vol. 
21
 (pg. 
674
-
687
)
McLachlan, A. and Jaramillo, E. 1995. Zonation on Sandy Beaches. Oceanography and Marine Biology: an Annual Review 33: 305-335.
Zonation on Sandy Beaches
Oceanography and Marine Biology: an Annual Review
, vol. 
33
 (pg. 
305
-
335
)
McLachlan, A., Jaramillo, E., Defeo, O., Dugan, J., de Ruyck, A. and Coetzee, P. 1995. Adaptations of bivalves to different beach types. Journal of Experimental Marine Biology and Ecology 187: 147-160.
Adaptations of bivalves to different beach types
Journal of Experimental Marine Biology and Ecology
, vol. 
187
 (pg. 
147
-
160
)
McLachlan, A., Jaramillo, E., Donn, T.E. and Wessels, F. 1993. Sandy beach macrofauna communities and their control by the physical environment: a geographical comparison. Journal of Coastal Research 15: 27-38.
Sandy beach macrofauna communities and their control by the physical environment: a geographical comparison
Journal of Coastal Research
, vol. 
15
 (pg. 
27
-
38
)
McLachlan, A., Wooldridge, T. and Dye, A.H. 1981. The ecology of sandy beaches in southern Africa. South African Journal of Zoology 16: 219-231.
The ecology of sandy beaches in southern Africa
South African Journal of Zoology
, vol. 
16
 (pg. 
219
-
231
)
Moore, P.G. and Francis, C.H. 1985. Water relations and osmoregulation of the beach-hopper Orchestia gammarellus (Pallas) (Crustacea: Amphipoda). Journal of Experimental Marine Biology and Ecology 94: 131-150.
Water relations and osmoregulation of the beach-hopper
Journal of Experimental Marine Biology and Ecology
, vol. 
94
 (pg. 
131
-
150
)
Mucci, A. 1983. The solubility of calcite and aragonite in seawater at various salinities, temperature, and one atmosphere total pressure. American Journal of Science 283: 780-799.
The solubility of calcite and aragonite in seawater at various salinities, temperature, and one atmosphere total pressure
American Journal of Science
, vol. 
283
 (pg. 
780
-
799
)
Nelson, W.G. (1993). Beach restoration in the southeastern US: environmental effects and biological monitoring. Ocean & Coastal Management 19: 157-182.
Beach restoration in the southeastern US: environmental effects and biological monitoring
Ocean & Coastal Management
, vol. 
19
 (pg. 
157
-
182
)
Nicholas, W. L and Trueman J.W.H. 2005. Biodiversity of marine nematodes in Australian sandy beaches from tropical and temperate regions. Biodiversity and Conservation 14: 823-839.
Biodiversity of marine nematodes in Australian sandy beaches from tropical and temperate regions
Biodiversity and Conservation
, vol. 
14
 (pg. 
823
-
839
)
Noy-Meir, I. 1979. Structure and function of desert ecosystems. Israel Journal of Botany 28: 1-19.
Structure and function of desert ecosystems
Israel Journal of Botany
, vol. 
78
 (pg. 
1
-
19
)
Oakeshott, R. 2003. Keynote speech presented to the 12th Annual New South Wales Coastal Conference, Port Macquarie, 4-7 November 2003.
O'Hara, T.D. 2002. Endemism, rarity and vulnerability of marine species along a temperate coastline. Invertebrate Systematics 16: 671-684.
Endemism, rarity and vulnerability of marine species along a temperate coastline
Invertebrate Systematics
, vol. 
16
 (pg. 
671
-
684
)
Pearse, A.S., Humm, H.J. and Wharton, G.W. 1942. Ecology of sandy beaches at Beaufort, North Carolina. Ecological Monographs 12: 135-190.
Ecology of sandy beaches at Beaufort, North Carolina
Ecological Monographs
, vol. 
12
 (pg. 
135
-
190
)
Peterson, C.H. and Bishop, M.J. 2005. Assessing the environmental impacts of beach nourishment. Bioscience 55: 887-896.
Assessing the environmental impacts of beach nourishment
Bioscience
, vol. 
55
 (pg. 
887
-
896
)
Peterson, C.H., Hickerson, D.H.M., Johnson, G.G. 2000. Short-term consequences of nourishment and bulldozing on the dominant large invertebrates of a sandy beach. Journal of Coastal Research 16: 368-378.
Short-term consequences of nourishment and bulldozing on the dominant large invertebrates of a sandy beach
Journal of Coastal Research
, vol. 
16
 (pg. 
368
-
378
)
Peterson, C.H., Bishop, M.J., Johnson, G.A., D'Anna, L. and Manning, L.M. 2006. Exploiting beach filling as an unaffordable experiment: upwards cascading impacts on sandy beach biota. Journal of Experimental Marine Biology and Ecology,doi:10.1016/j. jembe.2006.06.021.
Pörtner, H.O., Langenbuch, M. and Reipschläger, A. 2004. Biological impact of elevated ocean CO2 concentrations: lessons from animal physiology and earth history. Journal of Oceanography 60: 705-718.
Biological impact of elevated ocean CO2 concentrations: lessons from animal physiology and earth history
Journal of Oceanography
, vol. 
60
 (pg. 
705
-
718
)
Quammen, M.L. 1982. Influence of subtle substrate differences on feeding by shorebirds on intertidal mudflats. Marine Biology 71: 339-343.
Influence of subtle substrate differences on feeding by shorebirds on intertidal mudflats
Marine Biology
, vol. 
71
 (pg. 
339
-
343
)
Richardson, A.J. and Schoeman, D.S. 2004. Climate impact on plankton ecosystems in the Northeast Atlantic Science 305: 1609-1612.
Climate impact on plankton ecosystems in the Northeast Atlantic
Science
, vol. 
305
 (pg. 
1609
-
1612
)
Robertson, A.I. 1994. Sandy Beaches and Intertidal Flats. In: Marine Biology Eds L.S. Hammond and R.N. Synnot. Longman Cheshire Pty Ltd. Pp 297-314.
Schiel, D.R., Steinbeck, J.R. and Foster, M.S. 2004. Ten years of induced ocean warming causes comprehensive changes in marine benthic communities. Ecology 85: 1833-1839.
Ten years of induced ocean warming causes comprehensive changes in marine benthic communities
Ecology
, vol. 
85
 (pg. 
1833
-
1839
)
Schoeman, D.S. and Richardson, A.J. 2002. Investigating biotic and abiotic factors affecting the recruitment of an intertidal clam on an exposed sandy beach using a generalized additive model. Journal of Experimental Marine Biology and Ecology 276: 67-81.
Investigating biotic and abiotic factors affecting the recruitment of an intertidal clam on an exposed sandy beach using a generalized additive model
Journal of Experimental Marine Biology and Ecology
, vol. 
276
 (pg. 
67
-
81
)
Sherman, D.J., Barron, K.M. and Ellis, J.T. 2002. Retention of beach sands by dams and debris basins in Southern California. Journal of Coastal Research SI 36: 662-674.
Retention of beach sands by dams and debris basins in Southern California
Journal of Coastal Research
, vol. 
SI 36
 (pg. 
662
-
674
)
Sherman, K. and Duda, A.M. 1999. An ecosystem approach to global assessment and management of coastal waters. Marine Ecology Progress Series 190: 271-287.
An ecosystem approach to global assessment and management of coastal waters
Marine Ecology Progress Series
, vol. 
190
 (pg. 
271
-
287
)
Short, A.D. and Hesp, P.A. 1982. Wave, beach and dune interactions in southeastern Australia. Marine Geology 48: 259-284.
Wave, beach and dune interactions in southeastern Australia
Marine Geology
, vol. 
48
 (pg. 
259
-
284
)
Short, A.D. and McLeod, K. 1996. Australia's dynamic beaches: Nature, hazards and management. In: Proceedings of the Australian Coastal Management Conference (pp.200-204). Glenelg, Adelaide, South Australia: The University of Adelaide.
Proceedings of the Australian Coastal Management Conference
(pg. 
200
-
204
)
Short, A.D. and Wright, L.D. 1984. Morphodynamics of High Energy Beaches: an Australian Perspective. In: Coastal Geomorphology in Australia. Ed. B.G. Thom. Academic Press, Australia.
Coastal Geomorphology in Australia
Short, A.D., Trembanis, A. and Turner, I. 2000. Beach oscillation, rotation and the Southern Oscillation: Narrabeen Beach, Australia. Abstracts of the 27th International Conference on Coastal Engineering, July 16-22, 2000, Sydney, Australia. Vol. 1, Paper 4. Australian Institution of Engineers, Barton, ACT, Australia.
Soares, A.G., McLachlan, A. and Schlacher, T.A. 1996 Disturbance effects of stranded kelp on populations of the sandy beach bivalve Donax serra (Röding). Journal of Experimental Marine Biology and Ecology 205: 165-186.
Disturbance effects of stranded kelp on populations of the sandy beach bivalve Donax serra (Röding)
Journal of Experimental Marine Biology and Ecology
, vol. 
205
 (pg. 
165
-
186
)
Southward, A.J., Hawkins, S.J. and Burrows, M.T. 1995 Seventy years' observations of changes in distribution and abundance of zooplankton and intertidal organisms in the western English Channel in relation to rising sea temperature. Journal of Thermal Biology 20: 127-155.
Seventy years' observations of changes in distribution and abundance of zooplankton and intertidal organisms in the western English Channel in relation to rising sea temperature
Journal of Thermal Biology
, vol. 
20
 (pg. 
127
-
155
)
Speybroeck, J., Bonte, D., Courtens, W., Gheskiere, T., Grootaert, P., Maelfait, J-P., Mathys, M., Provoost, S., Sabbe, K., Stienen, W.M., Van Lancker, V., Vincx, M. and Degraer, S. 2006. Beach nourishment: an ecologically sound coastal defence alternative? A review. Aquatic Conservation: Marine and Freshwater Ecosystems 16: 419-435.
Beach nourishment: an ecologically sound coastal defence alternative? A review
Aquatic Conservation: Marine and Freshwater Ecosystems
, vol. 
16
 (pg. 
419
-
435
)
Taylor, J.D. and Reid, D.G. 1990. Shell microstructure and mineralogy of the Littorinidae: ecological and evolutionary significance. Hydrobiologia 193: 199-215.
Shell microstructure and mineralogy of the Littorinidae: ecological and evolutionary significance
Hydrobiologia
, vol. 
193
 (pg. 
199
-
215
)
Tomlinson, R. 2002. Beaches our asset. Planning and management for natural variability on open coastlines. In: Proceedings of the Public Workshop “Beach Protection and Management”. Yeppoon 2002. CRC for Coastal Zone, Estuary and Water Management and Faculty of Engineering and Physical Systems. Ed. J. Piorewisz.
Turley, C., Blackford, J., Widdicombe, S., Lowe, D. and Nightingale, P. 2005. Report presented at: International Symposium on the Stabilisation of Greenhouse Gas Concentrations. Exeter, UK. 1-3 February 2005.
Vermeij, G.J. 1979. Shell architecture and causes of death of Micronesian reef snails. Evolution 33: 686-696.
Shell architecture and causes of death of Micronesian reef snails
Evolution
, vol. 
33
 (pg. 
686
-
696
)
Von Koningsveld, M, Stive, M.J.F., Mulder, J.P.M, De Vriend, H.J., Ruessink, B.G. and Dunsbergen, D.W. 2003. Usefulness and effectiveness of coastal research: a matter of perception? Journal of Coastal Research 19: 441-461.
Usefulness and effectiveness of coastal research: a matter of perception?
Journal of Coastal Research
, vol. 
19
 (pg. 
441
-
461
)
Walther, G-R., Post, E., Convey, P., Menzels, A., Parmesan, C., Beebee, T.J.C., Fromentin, J-M., Hoegh-Guldberg, O. and Bairlein, F. 2002. Ecological responses to recent climate change. Nature 416: 389-395.
Ecological responses to recent climate change
Nature
, vol. 
416
 (pg. 
389
-
395
)
Weslawski J.M., Stanek, A., Siewart, A. and Beer, N. 2000. The sandhopper (Talitrus saltator, Montagu 1808) on the Polish Baltic coast. Is it a victim of increased tourism? Oceanological Studies 24: 77-87.
The sandhopper (Talitrus saltator, Montagu 1808) on the Polish Baltic coast. Is it a victim of increased tourism?
Oceanological Studies
, vol. 
24
 (pg. 
77
-
87
)
Wieser, W., Ott, J., Schiemer, F., and Gnaiger, E. 1974. An ecophysiological study of some meiofauna species inhabiting a sandy beach at Bermuda. Marine Biology 26: 235-248.
An ecophysiological study of some meiofauna species inhabiting a sandy beach at Bermuda
Marine Biology
, vol. 
26
 (pg. 
235
-
248
)
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