Bruxner Park Flora Reserve and the adjoining Ulidarra National Park, on the NSW north coast, comprise a dual-tenure conservation area of 1090 ha dominated by rainforest and old-growth moist eucalypt forest. A field survey of the area's vertebrate fauna over the period 1996-2005 detected 224 native species and nine introduced species. Incorporating records from secondary sources, the Bruxner-Ulidarra conservation area is known to support 230 native vertebrate species (39 mammals, 124 birds, 36 reptiles, 21 frogs and 10 fishes), including 20 species that are currently listed as threatened under NSW legislation. One hundred and ninety vertebrate species (79% of the total) were recorded during a systematic survey, with the remainder of records from opportunistic survey (18%) or secondary sources (3%). The effectiveness of the systematic survey effort in detecting the full inventory of species known from the study area ranged from 100% for freshwater fishes to only 61% for reptiles. Many reptile species were cryptic and/or at low density, and the resulting low detectability could easily have been mistaken for low reptile species diversity. This study demonstrates that a range of complementary survey methods over an extended period of time is needed to adequately sample the largest proportion of an area's total suite of vertebrate fauna. Suggestions are made concerning how to optimise results from vertebrate fauna inventory surveys.

Anderson, J.R. and Howland, M.B. 1998. Clarence/Coffs Harbour Regional Water Supply Project. Environmental Assessment - Working Paper No. 1: Aquatic Fauna. Unpublished report prepared for the Department of Public Works and Services.
Barker, J., Grigg, G.C. and Tyler, M.J. 1995. A Field Guide to Australian Frogs. Surrey Beatty and Sons, Chipping Norton, NSW.
A Field Guide to Australian Frogs
Barker, J., Lunney, D. and Bubela, T. 1994. Mammal surveys in the forests of the Carrai Plateau and Richmond Range in north-east New South Wales. Australian Mammalogy 17: 19-29.
Mammal surveys in the forests of the Carrai Plateau and Richmond Range in north-east New South Wales
Australian Mammalogy
, vol. 
17
 (pg. 
19
-
29
)
Bevan, J. 1983. A defensive reaction of Gonocephalus spinipes (Dumeril). Herpetofauna 14(2): 99.
A defensive reaction of Gonocephalus spinipes (Dumeril)
Herpetofauna
, vol. 
14
 pg. 
99
 
Cameron, R.A.D. and Pokryszko, B.M. 2005. Estimating the species richness and composition of land mollusc communities: problems, consequences and practical advice. Journal of Conchology 38(5): 529-547.
Estimating the species richness and composition of land mollusc communities: problems, consequences and practical advice
Journal of Conchology
, vol. 
38
 (pg. 
529
-
547
)
Cashner, R.C., Hawkes, G.P., Gartside, D.F. and Marsh-Matthews, E. 1999. Fishes of the Nymboida, Mann and Orara rivers of the Clarence river drainage, New South Wales, Australia. Proceedings of the Linnaean Society of NSW 121: 89-100.
Fishes of the Nymboida, Mann and Orara rivers of the Clarence river drainage, New South Wales, Australia
Proceedings of the Linnaean Society of NSW
, vol. 
121
 (pg. 
89
-
100
)
Catling, P.C., Burt, R.J. and Kooyman, R. 1997. A comparison of techniques used in a survey of the ground-dwelling and arboreal mammals in forests in north-eastern New South Wales. Wildlife Research 24: 417-32.
A comparison of techniques used in a survey of the ground-dwelling and arboreal mammals in forests in north-eastern New South Wales
Wildlife Research
, vol. 
24
 (pg. 
417
-
32
)
Churchill, S. 1998. Australian Bats. New Holland Publishers, Sydney, NSW.
Australian Bats
Christidis, L. and Boles, W. 1994. The Taxonomy and Species of Birds of Australia and its Territories. Royal Australian Ornithological Union Monograph 2.
Clemann, N., Long, K., Skurrie, D. and Dzuris, J. 2005. A trapping survey of small, ground-dwelling vertebrates in the Little Desert National Park, Victoria. Australian Zoologist 33: 119-126.
A trapping survey of small, ground-dwelling vertebrates in the Little Desert National Park, Victoria
Australian Zoologist
, vol. 
33
 (pg. 
119
-
126
)
Date, E.M., Ford, H.A. and Recher, H.F. 1991. Frugivorous pigeons, stepping stones, and weeds in northern New South Wales. Pp. 241-45 in Nature Conservation 2: The Role of Corridors, edited by D.A. Saunders and R.J. Hobbs. Surrey Beatty and Sons, Sydney, NSW.
Nature Conservation 2: The Role of Corridors
(pg. 
241
-
45
)
Debus, S.J.S. 1995. Surveys of large forest owls in northern NSW: methodology, calling behaviour and owl responses. Corella 19(2): 38-50.
Surveys of large forest owls in northern NSW: methodology, calling behaviour and owl responses
Corella
, vol. 
19
 (pg. 
38
-
50
)
Donnellan, S.C. and Mahony, M.J. 2004. Allozyme, chromosomal and morphological variability in the Litoria lesueuri species group (Anura: Hylidae), including a description of a new species. Australian Journal of Zoology 52: 1-28.
Allozyme, chromosomal and morphological variability in the Litoria lesueuri species group (Anura: Hylidae), including a description of a new species
Australian Journal of Zoology
, vol. 
52
 (pg. 
1
-
28
)
Donnellan, S.C., McGuigan, K., Knowles, R., Mahony, M.J. and Moritz, C. 1999. Genetic evidence for species boundaries in frogs of the Litoria citropa species-group (Anura: Hylidae). Australian Journal of Zoology 47: 275-93.
Genetic evidence for species boundaries in frogs of the Litoria citropa species-group (Anura: Hylidae)
Australian Journal of Zoology
, vol. 
47
 (pg. 
275
-
93
)
Dunsany, E.P. 1924. The King of Elfland's Daughter. G.P. Putnam and Sons, London.
The King of Elfland's Daughter
Ehmann, H. (editor) 1997. Threatened Frogs of New South Wales: Habitats, Status and Conservation. Frog and Tadpole Study Group of NSW Inc., Sydney, NSW.
Threatened Frogs of New South Wales: Habitats, Status and Conservation
Faragher, R.A. and Harris, J.H. 1994. The historical and current status of freshwater fish in New South Wales. Australian Zoologist 29: 166-176.
The historical and current status of freshwater fish in New South Wales
Australian Zoologist
, vol. 
29
 (pg. 
166
-
176
)
Fitzgerald, M., Shine, R. and Lemckert, F. 2002a. Radiotelemetric study of habitat use by the arboreal snake Hoplocephalus stephensii (Elapidae) in eastern Australia. Copeia 2002(2): 321-32.
Radiotelemetric study of habitat use by the arboreal snake Hoplocephalus stephensii (Elapidae) in eastern Australia
Copeia
, vol. 
2002
 (pg. 
321
-
32
)
Fitzgerald, M., Shine, R. and Lemckert, F. 2002b. Spatial ecology of arboreal snakes (Hoplocephalus stephensii, Elapidae) in an eastern Australian forest. Austral Ecology 27: 537-45.
Spatial ecology of arboreal snakes (Hoplocephalus stephensii, Elapidae) in an eastern Australian forest
Austral Ecology
, vol. 
27
 (pg. 
537
-
45
)
Fitzgerald, M., Shine, R. and Lemckert, F. 2004. Life history attributes of the threatened Australian snake (Stephens' banded snake Hoplocephalus stephensii, Elapidae). Biological Conservation 119: 121-28.
Life history attributes of the threatened Australian snake (Stephens' banded snake Hoplocephalus stephensii, Elapidae)
Biological Conservation
, vol. 
119
 (pg. 
121
-
28
)
Forestry Commission of New South Wales. 1989. Forest Preservation in State Forests of New South Wales. Research Note No. 47. Forestry Commission of NSW, Epping, NSW.
Forest Preservation in State Forests of New South Wales
Franklin, D.C. and Noske, R.A. 1998. Local movements of honeyeaters in a subcoastal vegetation mosaic in the Northern Territory. Corella 22: 97-103.
Local movements of honeyeaters in a subcoastal vegetation mosaic in the Northern Territory
Corella
, vol. 
22
 (pg. 
97
-
103
)
Garden, J.G., McAlpine, C.A., Possingham, H.P. and Jones, D.N. 2007. Using multiple survey methods to detect terrestrial reptiles and mammals: what are the most successful and cost-efficient combinations? Wildlife Research 34: 218-227.
Using multiple survey methods to detect terrestrial reptiles and mammals: what are the most successful and cost-efficient combinations
Wildlife Research
, vol. 
34
 (pg. 
218
-
227
)
Gibbons, P. and Lindenmayer, D. 2002. Tree Hollows and Wildlife Conservation in Australia. CSIRO Publishing, Collingwood, Victoria.
Tree Hollows and Wildlife Conservation in Australia
Gilmour, A. and Parnaby, H. 1994. Vertebrate Fauna of Conservation Concern in North-east NSW Forests. North East Forests Biodiversity Study Report No. 3e. Unpublished report, NSW National Parks and Wildlife Service, Sydney, NSW.
Kavanagh, R.P., Debus, S., Tweedie, T. and Webster, R. 1995. Distribution of nocturnal forest birds and mammals in northeastern New South Wales: relationships with environmental variables and management history. Wildlife Research 22: 359-377.
Distribution of nocturnal forest birds and mammals in northeastern New South Wales: relationships with environmental variables and management history
Wildlife Research
, vol. 
22
 (pg. 
359
-
377
)
Kemp, J.E. and Kutt, A.S. 2004. The vertebrate fauna of the Clemant State Forest lowlands: a significant coastal woodland remnant in the southern wet tropics bioregion, north-eastern Queensland. Australian Zoologist 32: 508-542.
The vertebrate fauna of the Clemant State Forest lowlands: a significant coastal woodland remnant in the southern wet tropics bioregion, north-eastern Queensland
Australian Zoologist
, vol. 
32
 (pg. 
508
-
542
)
Kikkawa, J. and Pearse, K. 1969. Geographical distribution of land birds in Australia - a numerical analysis. Australian Journal of Zoology 17: 821-40.
Geographical distribution of land birds in Australia - a numerical analysis
Australian Journal of Zoology
, vol. 
17
 (pg. 
821
-
40
)
Keogh, S.J., Scott, I.A.W., Fitzgerald, M. and Shine, R. 2003. Molecular phylogeny of the Australian venomous snake genus Hoplocephalus (Serpentes, Elapidae) and conservation genetics of the threatened H. stephensii. Conservation Genetics 4: 57-64.
Molecular phylogeny of the Australian venomous snake genus Hoplocephalus (Serpentes, Elapidae) and conservation genetics of the threatene
H. stephensii. Conservation Genetics
, vol. 
4
 (pg. 
57
-
64
)
Koehn, J.D. 1993. Fish need trees. The Victorian Naturalist 110(6): 255-257.
Fish need trees
The Victorian Naturalist
, vol. 
110
 (pg. 
255
-
257
)
Kutt, A.S., Kemp, J.E., McDonald, K.R., Williams, Y. , Williams, S.E., Hines, H.B., Hero, J-M. and Torr, G. 2005. Vertebrate fauna survey of White Mountains National Park in the desert uplands bioregion, central-north Queensland. Australian Zoologist 33: 17-34.
Vertebrate fauna survey of White Mountains National Park in the desert uplands bioregion, central-north Queensland
Australian Zoologist
, vol. 
33
 (pg. 
17
-
34
)
Lloyd, A. 2004. Bat distribution and riparian zone management in production forests of north-eastern New South Wales. Unpublished Honours thesis. Southern Cross University, Lismore, NSW.
Lunney, D. and Barker, J. 1986. Survey of reptiles and amphibians of the coastal forests near Bega, NSW. Australian Zoologist 22: 1-9.
Survey of reptiles and amphibians of the coastal forests near Bega, NSW
Australian Zoologist
, vol. 
22
 (pg. 
1
-
9
)
Lunney, D., Matthews, A., Cogger, H. and Dickman, C. 2004. The neglected 74% - the non-threatened vertebrates - and a reflection on the limitations of the process that fashioned the current schedules of threatened species in New South Wales. Pp. 145-157 in Threatened Species Legislation: is it just an Act?, edited by P. Hutchings, D. Lunney and C. Dickman. Royal Zoological Society of New South Wales, Mosman, NSW.
Threatened Species Legislation: is it just an Act?
(pg. 
145
-
157
)
Mahony, M.J. 1993. The status of frogs in the Watagan Mountains area the central coast of New South Wales. Pp. 257-264 in Herpetology in Australia: A Diverse Discipline, edited by D. Lunney and D. Ayers, Royal Zoological Society of New South Wales, Mosman, NSW.
Herpetology in Australia: A Diverse Discipline
(pg. 
257
-
264
)
McDowell, R. (editor). 1996. Freshwater Fishes of South-eastern Australia. Reed Books, Chatswood, NSW.
Freshwater Fishes of South-eastern Australia
Menkhorst, P. and Knight, F. 2001. A Field Guide to the Mammals of Australia. Oxford University Press, Melbourne, Victoria.
A Field Guide to the Mammals of Australia
Milledge, D. 1991. A survey of the terrestrial vertebrates of coastal Byron Shire. Australian Zoologist 27: 66-90.
A survey of the terrestrial vertebrates of coastal Byron Shire
Australian Zoologist
, vol. 
27
 (pg. 
66
-
90
)
Milton, D.A. 1980. A comparison of three techniques used in a reptile survey of the Conondale Ranges. The Victorian Naturalist 97(1): 26-31.
A comparison of three techniques used in a reptile survey of the Conondale Ranges
The Victorian Naturalist
, vol. 
97
 (pg. 
26
-
31
)
Moran, C., Catterall, C.P., Green, R.J. and Olsen, M.F. 2004. Fates of feathered fruit-eaters in fragmented forests. Pp. 699-712 in Conservation of Australia's Forest Fauna (2nd edition), edited by D. Lunney. Royal Zoological Society of New South Wales, Mosman, NSW.
Conservation of Australia's Forest Fauna
(pg. 
699
-
712
)
Morris, S.A., Pollard, D.A., Gehrke, P.C. and Pogonoski, J.J. 2001. Threatened and Potentially Threatened Freshwater Fishes of Coastal New South Wales and the Murray-Darling Basin. NSW Fisheries Final Report Series No. 33. NSW Fisheries, Sydney, NSW.
Threatened and Potentially Threatened Freshwater Fishes of Coastal New South Wales and the Murray-Darling Basin
Murphy, M.J. 2007. The diverse land snail community of Bruxner Park on the north coast of New South Wales, Australia. The Victorian Naturalist 124(5): 306-309.
The diverse land snail community of Bruxner Park on the north coast of New South Wales, Australia
The Victorian Naturalist
, vol. 
124
 (pg. 
306
-
309
)
Musyl, M.K. and Keenan, C.P. 1996. Evidence for cryptic speciation in Australian freshwater eel-tailed catfish, Tandanus tandanus (Teleostei: Plotosidae). Copeia 1996(3): 526-534.
Evidence for cryptic speciation in Australian freshwater eel-tailed catfish, Tandanus tandanus (Teleostei: Plotosidae)
Copeia
, vol. 
1996
 (pg. 
526
-
534
)
Parris, K.M., Norton, T. W. and Cunningham, R.B. 1999. A comparison of techniques for sampling amphibian communities in the forests of southeast Queensland, Australia. Herpetologica 55: 271-83.
A comparison of techniques for sampling amphibian communities in the forests of southeast Queensland, Australia
Herpetologica
, vol. 
55
 (pg. 
271
-
83
)
Roberts, P.E. 1971. Bruxner Park Flora Reserve Preliminary Bird List. Unpublished report. Copy held by Forests NSW, Coffs Harbour, NSW.
Schilthuizen, M. and Rutjes, H.A. 2001. Land snail diversity in a square kilometre of tropical rainforest in Sabah, Malaysian Borneo. Journal of Molluscan Studies 67: 417-423.
Land snail diversity in a square kilometre of tropical rainforest in Sabah, Malaysian Borneo
Journal of Molluscan Studies
, vol. 
67
 (pg. 
417
-
423
)
Schodde, R. 1979. The mysterious origins of Australian birds. Pp. 602-05 in Reader's Digest Complete Book of Australian Birds, edited by H. J. Frith. Reader's Digest Services, Sydney, NSW.
Reader's Digest Complete Book of Australian Birds
(pg. 
602
-
05
)
Schulz, M. 1993. Reptiles and amphibians of the coastal dunes at Venus Bay, Victoria. The Victorian Naturalist 110(5): 198-204.
Reptiles and amphibians of the coastal dunes at Venus Bay, Victoria
The Victorian Naturalist
, vol. 
110
 (pg. 
198
-
204
)
Schulz, M. and de Oliveira, M.C. 1995. Microchiropteran fauna of Kroombit Tops, central Queensland, including a discussion on survey techniques. Australian Zoologist 30: 71-77.
Microchiropteran fauna of Kroombit Tops, central Queensland, including a discussion on survey techniques
Australian Zoologist
, vol. 
30
 (pg. 
71
-
77
)
Scotts, D. 1991. Old-growth forests: their ecological characteristics and value to forest-dependent vertebrate fauna of south-east Australia. Pp. 147-59 in Conservation of Australia's Forest Fauna, edited by D. Lunney. Royal Zoological Society of New South Wales, Mosman, NSW.
Conservation of Australia's Forest Fauna
(pg. 
147
-
59
)
Spector, S. 2002. Biogeographic crossroads as priority areas for biodiversity conservation. Conservation Biology 16: 1480-1487.
Biogeographic crossroads as priority areas for biodiversity conservation
Conservation Biology
, vol. 
16
 (pg. 
1480
-
1487
)
State of the Environment Advisory Council. 1996. Australian State of the Environment 1996. CSIRO Publishing, Collingwood, Victoria.
Australian State of the Environment 1996
Swan, G., Shea, G. and Sadlier, R. 2004. A Field Guide to Reptiles of New South Wales. 2nd Edition. Reed New Holland, Sydney, NSW.
A Field Guide to Reptiles of New South Wales
Tasker, E.M. And Dickman, C.R. 2002. A review of Elliott trapping methods for small mammals in Australia. Australian Mammalogy 23: 77-87.
A review of Elliott trapping methods for small mammals in Australia
Australian Mammalogy
, vol. 
23
 (pg. 
77
-
87
)
Thackway, R. and Creswell, I.D. (editors) 1995. An Interim Biogeographic Regionalisation for Australia: A Framework for Establishing the National System of Reserves Version 4.0. Australian Nature Conservation Agency, Canberra, ACT.
An Interim Biogeographic Regionalisation for Australia: A Framework for Establishing the National System of Reserves
Webb, L.J., Tracey, J.G. and Williams, W. T. 1984. A floristic framework of Australian rainforests. Australian Journal of Ecology 9: 169-198.
A floristic framework of Australian rainforests
Australian Journal of Ecology
, vol. 
9
 (pg. 
169
-
198
)
Woinarski, J.C.Z., Armstrong, M., Brennan, K., Connors, G., Milne, D., McKenzie, G. and Edwards, K. 2000. A different fauna?: captures of vertebrates in a pipeline trench, compared with conventional survey techniques; and a consideration of mortality patterns in a pipeline trench. Australian Zoologist: 31: 421-431.
A different fauna?: captures of vertebrates in a pipeline trench, compared with conventional survey techniques; and a consideration of mortality patterns in a pipeline trench
Australian Zoologist
, vol. 
31
 (pg. 
421
-
431
)
Wood, D.H. 1971. The ecology of Rattus fuscipes and Melomys cervinipes (Rodentia: Muridae) in a south-east Queensland rain forest. Australian Journal of Zoology 19: 371-392.
The ecology of Rattus fuscipes and Melomys cervinipes (Rodentia: Muridae) in a south-east Queensland rain forest
Australian Journal of Zoology
, vol. 
19
 (pg. 
371
-
392
)
Clark, S. 2009. A review of the land snail genus Meridolum (Gastropoda: Camaenidae) from central New South Wales, Australia. Molluscan Research 29(2): 61-120.
A review of the land snail genus Meridolum (Gastropoda: Camaenidae) from central New South Wales, Australia
Molluscan Research
, vol. 
29
 (pg. 
61
-
120
)
Couper, P.J., Sadlier, R.A., Shea, G.M. and Worthington Wilmer, J. 2008. A reassessment of Saltuarius swaini (Lacertilia: Diplodactylidae) in southeastern Queensland and New South Wales; two new taxa, phylogeny, biogeography and conservation. Records of the Australian Museum 60: 87-118.
A reassessment of Saltuarius swaini (Lacertilia: Diplodactylidae) in southeastern Queensland and New South Wales; two new taxa, phylogeny, biogeography and conservation
Records of the Australian Museum
, vol. 
60
 (pg. 
87
-
118
)
Murphy, M.J. in press. Notes on the butterflies of Bruxner Park on the north coast of New South Wales, Australia. The Victorian Naturalist.
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