The koala Phascolarctos cinereus is the only member of the once diverse marsupial family Phascolarctidae to have survived the Last Glacial Maximum. A climate envelope model for P. cinereus was developed to predict the range for this species at present and at the Last Glacial Maximum. The model was compared to the contemporary koala records and the known fossil records of P. cinereus during the Quaternary. The predicted current core range for koalas was concentrated in southeast Queensland,eastern New South Wales and eastern Victoria. At the Last Glacial Maximum their predicted core range contracted significantly to southeast Queensland and northeast New South Wales. Our findings concord with other studies that find species experienced range contractions during glacial maxima. In the context of the future conservation planning for koalas in the wild, our historical perspective demonstrates the past adaptations of koalas to changes in climate and their probable range contraction to climatic refugia. The future survival of wide-ranging specialist species, such as the koala, may depend on identifying and protecting, future climatic refugia.

Climate envelope, climatic refugia, core range, koala, Phascolarctos cinereus, Last Glacial Maximum
Allen, C. D. and Breshears, D. D. 1998. Droughtinduced shift of a forest-woodland ecotone: rapid landscape response to climate variation. Proceedings of the National Academy of Sciences of the United States of America 95: 14839-14842.
,
Droughtinduced shift of a forest-woodland ecotone: rapid landscape response to climate variation
,
Proceedings of the National Academy of Sciences of the United States of America
, vol.
95
(pg.
14839
-
14842
)
Archer, M. 1972. Phascolarctos (Marsupialia, Vombatoidea) and associated fossil fauna from Koala Cave near Yanchep, Western Australia. Helictite 49-59.
,
Phascolarctos (Marsupialia, Vombatoidea) and associated fossil fauna from Koala Cave near Yanchep, Western Australia
,
Helictite
(pg.
45
-
59
)
Archer, M., Hand, S.J., and Godthelp, H. 2000. Australia's Lost World: Riversleigh, World Heritage Site. Reed New Holland, Australia.
,
Australia's Lost World: Riversleigh, World Heritage Site
Ashcroft, M.B. 2010. Identifying refugia from climate change. Journal of Biogeography 37: 1407-1413.
,
Identifying refugia from climate change
,
Journal of Biogeography
, vol.
37
(pg.
1407
-
1413
)
Balme, L.M., Merrilees, D. and Porter, J.K. 1978. Late Quaternary mammal remains spanning 30,000 years from excavations in Devil's Lair, Western Australia. Journal of the Royal Society of Western Australia 61: 33-65.
,
Late Quaternary mammal remains spanning 30,000 years from excavations in Devil's Lair, Western Australia
,
Journal of the Royal Society of Western Australia
, vol.
61
(pg.
33
-
65
)
Barrows, T. T., Juggins, S., De Deckker, P., Calvo, E. and Pelejero, C. 2007. Long-term sea surface temperature and climate change in the Australian-New Zealand region. Paleoceanography 22, 2: Article number PA2215.
Bartholomai, A. 1977. The fossil vertebrate fauna from Pleistocene deposits at Cement Mills, Gore, southeastern Queensland. Memoirs of the Queensland Museum 18: 41-51.
,
The fossil vertebrate fauna from Pleistocene deposits at Cement Mills, Gore, southeastern Queensland
,
Memoirs of the Queensland Museum
, vol.
18
(pg.
41
-
51
)
Beaumont, L. J., Hughes, L. and Poulsen, M. 2005. Predicting species distributions: use of climatic parameters in BIOCLIM and its impact on predictions of species' current and future distributions. Ecological Modelling 186: 250-269.
,
Predicting species distributions: use of climatic parameters in BIOCLIM and its impact on predictions of species' current and future distributions
,
Ecological Modelling
, vol.
186
(pg.
250
-
269
)
Black, K. 1999. Diversity and relationships of living and extinct koalas (Phascolarctidae, Marsupialia). Australian Mammalogy 21: 16-17.
,
Diversity and relationships of living and extinct koalas (Phascolarctidae, Marsupialia)
,
Australian Mammalogy
, vol.
21
(pg.
16
-
17
)
Box, E. O. 1981. Predicting physiognomic vegetation types with climate variables. Plant Ecology 45: 127-39.
,
Predicting physiognomic vegetation types with climate variables
,
Plant Ecology
, vol.
45
(pg.
127
-
39
)
Cunningham, S.C., Thomson, J.R., Read, J., Baker, P.J. and Nally, R.M. 2010. Does stand structure influence susceptibility of eucalypt floodplain forests to dieback? Austral Ecology 35: 348-356.
,
Does stand structure influence susceptibility of eucalypt floodplain forests to dieback
,
Austral Ecology
, vol.
35
(pg.
348
-
356
)
DeGabriel, J., Moore, B., Marsh, K. and Foley, W. 2010. The effect of plant secondary metabolites on the interplay between the internal and external environments of marsupial folivores. Chemoecology 20:97-108.
,
The effect of plant secondary metabolites on the interplay between the internal and external environments of marsupial folivores
,
Chemoecology
, vol.
20
(pg.
97
-
108
)
Dique, D.S., Thompson, J., Preece, H.J., Penfold, G.C., de Villiers, D.L. and Leslie, R.S. 2003. Koala mortality on roads in south-east Queensland: the koala speed-zone trial Wildlife Research 30: 419 – 426.
,
Koala mortality on roads in south-east Queensland: the koala speed-zone trial
,
Wildlife Research
, vol.
30
(pg.
419
-
426
)
Duka, T. and Masters, P. 2005. Confronting a tough issue: fertility control and translocation for over-abundant koalas on Kangaroo Island, South Australia. Ecological Management & Restoration 6: 172-81.
,
Confronting a tough issue: fertility control and translocation for over-abundant koalas on Kangaroo Island, South Australia
,
Ecological Management & Restoration
, vol.
6
(pg.
172
-
81
)
Ellis, W.A.H., Melzer, A., Carrick, F.N. and Hasegawa, M. 2002. Tree use, diet and home range of the koala (Phascolarctos cinereus) at Blair Athol, central Queensland. Wildlife Research 29: 303-311.
,
Tree use, diet and home range of the koala (Phascolarctos cinereus) at Blair Athol, central Queensland
,
Wildlife Research
, vol.
29
(pg.
303
-
311
)
Fensham, R. J. and Fairfax, R. J. 2007. Drought-related tree death of savanna eucalypts: species susceptibility, soil conditions and root architecture. Journal of Vegetation Science 18: 71-80.
,
Drought-related tree death of savanna eucalypts: species susceptibility, soil conditions and root architecture
,
Journal of Vegetation Science
, vol.
18
(pg.
71
-
80
)
Fowler, G. 1993. 'Black August' Queensland's Open Season on Koalas in 1927. Honours Thesis, Australian National University.
,
'Black August' Queensland's Open Season on Koalas in 1927
Gallagher, S. J., Greenwood, D. R., Taylor, D., Smith, A. J., Wallace, M. W. and Holdgate, G. R. 2003. The Pliocene climatic and environmental evolution of southeastern Australia: evidence from the marine and terrestrial realm. Palaeogeography, Palaeoclimatology, Palaeoecology 193: 349-382.
,
The Pliocene climatic and environmental evolution of southeastern Australia: evidence from the marine and terrestrial realm
,
Palaeogeography, Palaeoclimatology, Palaeoecology
, vol.
193
(pg.
349
-
382
)
Glauret, L. 1910. The Mammoth Cave. Records of the Western Australian Museum and Art Gallery 1: 11-36.
,
The Mammoth Cave
,
Records of the Western Australian Museum and Art Gallery
, vol.
1
(pg.
11
-
36
)
Gordon, G., Brown, A.S. and Pulsford, T. 1988. A koala (Phascolarctos cinereus Goldfuss) population crash during drought and heatwave conditions in south-western Queensland. Austral Ecology 13: 451-461.
,
A koala (Phascolarctos cinereus Goldfuss) population crash during drought and heatwave conditions in south-western Queensland
,
Austral Ecology
, vol.
13
(pg.
451
-
461
)
Green, K., Stein, J. A. and Driessen, M. M. 2008. The projected distributions of Mastacomys fuscus and Rattus lutreolus in south-eastern Australia under a scenario of climate change: potential for increased competition? Wildlife Research 35: 113-9.
,
The projected distributions of Mastacomys fuscus and Rattus lutreolus in south-eastern Australia under a scenario of climate change: potential for increased competition
,
Wildlife Research
, vol.
35
(pg.
113
-
9
)
Heikkinen, R. K., Luoto, M., Leikola, N., Poyry, J., Settele, J., Kudrna, O., Marmion, M., Fronzek, S. and Thuiller, W. 2009. Assessing the vulnerability of European butterflies to climate change using multiple criteria. Biodiversity and Conservation 19: 695-723.
,
Assessing the vulnerability of European butterflies to climate change using multiple criteria
,
Biodiversity and Conservation
, vol.
19
(pg.
695
-
723
)
Hernandez, P. A., Graham, C. H., Master, L. L. and Albert, D. L. 2006. The effect of sample size and species characteristics on performance of different species distribution modeling methods. Ecography 29: 773-85.
,
The effect of sample size and species characteristics on performance of different species distribution modeling methods
,
Ecography
, vol.
29
(pg.
773
-
85
)
Hewitt, G. 2000. The genetic legacy of the Quaternary ice ages. Nature 405: 907-913.
,
The genetic legacy of the Quaternary ice ages
,
Nature
, vol.
405
(pg.
907
-
913
)
Hijmans, R. J. and Graham, C. H. 2006. The ability of climate envelope models to predict the effect of climate change on species distributions. Global Change Biology 12: 2272-2281.
,
The ability of climate envelope models to predict the effect of climate change on species distributions
,
Global Change Biology
, vol.
12
(pg.
2272
-
2281
)
Hocknull, S. A., Zhao, J.-x., Feng, Y.-x. and Webb, G. E. 2007. Responses of Quaternary rainforest vertebrates to climate change in Australia. Earth and Planetary Science Letters, 264: 317-31.
,
Responses of Quaternary rainforest vertebrates to climate change in Australia
,
Earth and Planetary Science Letters
, vol.
264
(pg.
317
-
31
)
Hoegh-Guldberg, O., Hughes, L., McIntyre, S., Lindenmayer, D. B., Parmesan, C., Possingham, H. P. and Thomas, C. D. 2008. Assisted colonization and rapid climate change. Science 321: 345-346.
,
Assisted colonization and rapid climate change
,
Science
, vol.
321
(pg.
345
-
346
)
Houlder, D., Hutchinson, M., Nix, H. and McMahon, J. 1999. ANUCLIM Version 5.0 User Guide. The Australian National University, Centre for Resource and Environmental Studies, Canberra.
,
ANUCLIM Version 5.0 User Guide
Hrdina, F., and Gordon, G. 2004. The koala and possum trade in Queensland, 1906-1936. Australian Zoologist 32: 543-584.
,
The koala and possum trade in Queensland, 1906-1936
,
Australian Zoologist
, vol.
32
(pg.
543
-
584
)
Hughes, L., Cawsey, E. M. and Westoby, M. 1996. Climatic range sizes of Eucalyptus species in relation to future climate change. Global Ecology and Biogeography Letters 5: 23-29.
,
Climatic range sizes of Eucalyptus species in relation to future climate change
,
Global Ecology and Biogeography Letters
, vol.
5
(pg.
23
-
29
)
Jackson, S. 2007. Koala Origins of an Icon. Jacana Books, an imprint of Allen & Unwin, Crow's Nest, N.S.W. Australia.
Jackson, M., White, N., Giffard, P. and Timms, P. 1999. Epizootiology of Chlamydia infections in two freerange koala populations. Veterinary Microbiology 65: 225-234.
,
Epizootiology of Chlamydia infections in two freerange koala populations
,
Veterinary Microbiology
, vol.
65
(pg.
225
-
234
)
Johnson, C. 2006. Australia's Mammal Extinctions A 50 000 year history. Cambridge University Press, New York.
,
Australia's Mammal Extinctions A 50 000 year history
Klicka, J. and Zink, R. M. 1997. The importance of recent ice ages in speciation: a failed paradigm. Science 277: 1666-1669.
,
The importance of recent ice ages in speciation: a failed paradigm
,
Science
, vol.
277
(pg.
1666
-
1669
)
Knapp, S. and Mallet, J. 2003. Refuting refugia? Science 300: 71-72.
,
Refuting refugia
,
Science
, vol.
300
(pg.
71
-
72
)
Korsten, M., Ho, S.Y.W., Davison, J., Pahn, B., Vulla, E., Roht, M., Tumanov, I.L., Kojola, I., Andersone- Lilley, Z., Ozolins, J., Pilot, M., Mertzanis, Y., Giannakopoulos, A., Vorobiev, A.A., Markov, N.I., Saveljev, A.P., Lyapunova, E.A., Abramov, A.V., Mannil, P., Valdmann, H., Pazetnov, S.V., Pazetnov, V.S., Rokov, A.M. and Saarma, U. 2009. Sudden expansion of a single brown bear maternal lineage across northern continental Eurasia after the last ice age: A general demographic model for mammals? Molecular Ecology 18: 1963-1979.
,
Sudden expansion of a single brown bear maternal lineage across northern continental Eurasia after the last ice age: A general demographic model for mammals
,
Molecular Ecology
, vol.
18
(pg.
1963
-
1979
)
Krockenberger, A. 2003. Meeting the energy demands of reproduction in female koalas, Phascolarctos cinereus: evidence for energetic compensation. Journal of Comparative Physiology B-Biochemical Systemic and Environmental Physiology 173: 531-40.
,
Meeting the energy demands of reproduction in female koalas, Phascolarctos cinereus: evidence for energetic compensation
,
Journal of Comparative Physiology B-Biochemical Systemic and Environmental Physiology
, vol.
173
(pg.
531
-
40
)
Long, J., Archer, M., Flannery, T. and Hand, S. 2002. Prehistoric Mammals of Australia and New Guinea One Hundred Million Years of Evolution. University of New South Wales Press Limited, NSW.
,
Prehistoric Mammals of Australia and New Guinea One Hundred Million Years of Evolution
Louys, J., Aplin, K., Beck, R.M.D., and Archer, M. 2009. Cranial anatomy of Oligo-Miocene koalas (diprotodontia: phascolarctidae): stages in the evolution of an extreme leaf-eating specialization.
Lundelius Jr., E.L. and Turnbull, W.D. 1982. The mammalian fauna of Madura Cave, Western Australia. Part V: Diprotodonta (Part). Fieldiana: Geology 11: 1-31.
,
The mammalian fauna of Madura Cave, Western Australia
,
Part V: Diprotodonta (Part). Fieldiana: Geology
, vol.
11
(pg.
1
-
31
)
Lunney, D., Crowther, M.S., Shannon, I., and Bryant, J. V. 2009. Combining a map-based public survey with an estimation of site occupancy to determine the recent and changing distribution of the koala in New South Wales. Wildlife Research 36: 262–273.
,
Combining a map-based public survey with an estimation of site occupancy to determine the recent and changing distribution of the koala in New South Wales
,
Wildlife Research
, vol.
36
(pg.
262
-
273
)
Manning, A.D., Lindenmayer, D.B., Nix, H.A. and Barry, S.C. 2005. A bioclimatic analysis for the highly mobile Superb Parrot of south-eastern Australia. Emu 105: 193-201.
,
A bioclimatic analysis for the highly mobile Superb Parrot of south-eastern Australia
,
Emu
, vol.
105
(pg.
193
-
201
)
Marshall, L.G. 1973. Fossil vertebrate faunas from the Lake Victoria region, S.W. New South Wales, Australia. Memoirs of the National Museum of Victoria 34: 151-172.
,
Fossil vertebrate faunas from the Lake Victoria region, S.W. New South Wales, Australia
,
Memoirs of the National Museum of Victoria
, vol.
34
(pg.
151
-
172
)
McAlpine, C.A., Rhodes, J.R., Callaghan, J.G., Bowen, M.E., Lunney, D., Mitchell, D.L., Pullar, D.V. and Possingham, H.P. 2006. The importance of forest area and configuration relative to local habitat factors for conserving forest mammals: A case study of koalas in Queensland, Australia. Biological Conservation 132: 153-165.
,
The importance of forest area and configuration relative to local habitat factors for conserving forest mammals: A case study of koalas in Queensland, Australia
,
Biological Conservation
, vol.
132
(pg.
153
-
165
)
McKenney, D. W., Pedlar, J. H., Lawrence, K., Campbell, K. and Hutchinson, M. F. 2007. Beyond traditional hardiness zones: using climate envelopes to map plant range limits. BioScience 57: 929-37.
,
Beyond traditional hardiness zones: using climate envelopes to map plant range limits
,
BioScience
, vol.
57
(pg.
929
-
37
)
McLachlan, J. S., Hellmann, J. J. and Schwartz, M. W. 2007. A framework for debate of assisted migration in an era of climate change. Conservation Biology 21: 297-302.
,
A framework for debate of assisted migration in an era of climate change
,
Conservation Biology
, vol.
21
(pg.
297
-
302
)
Merrilees, D, 1967. Man the destroyer: late Quaternary changes in the Australian marsupial fauna. Journal of the Royal Society of Western Australia 51: 1-24.
,
Man the destroyer: late Quaternary changes in the Australian marsupial fauna
,
Journal of the Royal Society of Western Australia
, vol.
51
(pg.
1
-
24
)
Mesquita, S. and Sousa, A. J. 2009. Bioclimatic mapping using geostatistical approaches: application to mainland Portugal. International Journal of Climatology 29: 2156-2170.
,
Bioclimatic mapping using geostatistical approaches: application to mainland Portugal
,
International Journal of Climatology
, vol.
29
(pg.
2156
-
2170
)
Moore, B. D. and Foley, W. J. 2000. A review of feeding and diet selection in koalas (Phascolarctos cinereus). Australian Journal of Zoology 48: 317-33.
,
A review of feeding and diet selection in koalas (Phascolarctos cinereus)
,
Australian Journal of Zoology
, vol.
48
(pg.
317
-
33
)
Moore, B.D., Foley, W.J., Wallis, I.R., Cowling, A. and Handasyde, K.A. 2005. Eucalyptus foliar chemistry explains selective feeding by koalas. Biology Letters 1: 64-67.
,
Eucalyptus foliar chemistry explains selective feeding by koalas
,
Biology Letters
, vol.
1
(pg.
64
-
67
)
Moss, P. T. and Kershaw, P. A. 2000. The last glacial cycle from the humid tropics of northeastern Australia: comparison of a terrestrial and a marine record Palaeogeography, Palaeoclimatology, Palaeoecology 155: 155-176.
,
The last glacial cycle from the humid tropics of northeastern Australia: comparison of a terrestrial and a marine record
,
Palaeogeography, Palaeoclimatology, Palaeoecology
, vol.
155
(pg.
155
-
176
)
Nevill, P. G., Bossinger, G. and Ades, P. K. 2009. Phylogeography of the world's tallest angiosperm, Eucalyptus regnans: evidence for multiple isolated Quaternary refugia. Journal of Biogeography 37: 179-192.
,
Phylogeography of the world's tallest angiosperm, Eucalyptus regnans: evidence for multiple isolated Quaternary refugia
,
Journal of Biogeography
, vol.
37
(pg.
179
-
192
)
Petherick, L., McGowan, H. and Moss, P. 2008. Climate variability during the Last Glacial Maximum in eastern Australia: evidence of two stadials? Journal of Quaternary Science Published Online. DOI: 10.1002/JQS.1186.
Piper, K.J. 2007. Early Pleistocene mammals from the Nelson Bay local fauna, Portland, Victoria, Australia. Journal of Vertebrate Paleontology 27: 492-503.
,
Early Pleistocene mammals from the Nelson Bay local fauna, Portland, Victoria, Australia
,
Journal of Vertebrate Paleontology
, vol.
27
(pg.
492
-
503
)
Pledge, N.S. 1992. The Curramulka local fauna: a new late tertiary fossil assemblage from Yorke Peninsula, South Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 9: 115-142.
,
The Curramulka local fauna: a new late tertiary fossil assemblage from Yorke Peninsula, South Australia
,
The Beagle, Records of the Northern Territory Museum of Arts and Sciences
, vol.
9
(pg.
115
-
142
)
Price, G.J. 2008. Is the modern koala (Phascolarctos cinereus) a derived dwarf of a Pleistocene giant? Implications for testing megafauna extinction hypothesis. Quaternary Science Reviews 27: 2516-2521.
,
Is the modern koala (Phascolarctos cinereus) a derived dwarf of a Pleistocene giant
,
Implications for testing megafauna extinction hypothesis. Quaternary Science Reviews
, vol.
27
(pg.
2516
-
2521
)
Price, G. J., Zhao, J.-x., Feng, Y.-x. and Hocknull, S. A. 2009. New records of Plio-Pleistocene koalas from Australia: palaeoecological and taxonomic implications. Records of the Australian Museum 61: 39-48.
,
New records of Plio-Pleistocene koalas from Australia: palaeoecological and taxonomic implications
,
Records of the Australian Museum
, vol.
61
(pg.
39
-
48
)
Prideaux, G.J., Roberts, R.G., Megirian, D., Westaway, K.E., Hekkstrom, J.C. and Olley, J.M. 2007. Mammalian responses to Pleistocene climate change in southeastern Australia. Geology 35: 33-36.
,
Mammalian responses to Pleistocene climate change in southeastern Australia
,
Geology
, vol.
35
(pg.
33
-
36
)
Rhodes, J. R., Callaghan, J. G., McAlpine, C. A., De Jong, C., Bowen, M. E., Mitchell, D. L., Lunney, D. and Possingham, H. P. 2008. Regional variation in habitat-occupancy thresholds: a warning for conservation planning. Journal of Applied Ecology 45: 549-557.
,
Regional variation in habitat-occupancy thresholds: a warning for conservation planning
,
Journal of Applied Ecology
, vol.
45
(pg.
549
-
557
)
Sommer, R. S. and Zachos, F. E. 2009. Fossil evidence and phylogeography of temperate species: 'glacial refugia' and post-glacial recolonization. Journal of Biogeography 36: 2013-2020.
,
Fossil evidence and phylogeography of temperate species: 'glacial refugia' and post-glacial recolonization
,
Journal of Biogeography
, vol.
36
(pg.
2013
-
2020
)
State of Queensland 2009. Decline of the Koala Coast Koala Population: Population Status in 2008. Department of Environment and Resource Management, Queensland, Australia. http://www.derm.qld.gov.au/register/p02966aa.pdf
Stirton, R.A. 1957. A new koala from the Pliocene Palankarinna Fauna of South Australia. Records of the South Australian Museum 13: 71-81.
,
A new koala from the Pliocene Palankarinna Fauna of South Australia
,
Records of the South Australian Museum
, vol.
13
(pg.
71
-
81
)
Svenning, J., Normand, S. and Kageyama, M. 2008. Glacial refugia of temperate trees in Europe: insights from species distribution modelling. Journal of Ecology 96: 1117-27.
,
Glacial refugia of temperate trees in Europe: insights from species distribution modelling
,
Journal of Ecology
, vol.
96
(pg.
1117
-
27
)
Turney, C.S.M., Haberle, S., Fink, D., Kershaw, A.P., Barbetti, M., Barrows, T.T., Black, M., Cohen, T.J., Correge, T., Hesse, P.P., Hua, Q., Johnston, R., Morgan, V., Moss, P., Nanson, G., Van Ommen, T., Rule, S., Williams, N.J., Zhao, J.-X., D'Costa, D., Feng, Y.-X., Gagan, M., Mooney, S. and Xia, Q. 2006. Integration of ice-core, marine and terrestrial records for the Australian Last Glacial Maximum and Termination: a contribution from the OZ INTIMATE group. Journal of Quaternary Science 21: 751-761.
,
Integration of ice-core, marine and terrestrial records for the Australian Last Glacial Maximum and Termination: a contribution from the OZ INTIMATE group
,
Journal of Quaternary Science
, vol.
21
(pg.
751
-
761
)
Van der Kaars, S., De Deckker, P. and Gingele, F. X. 2006. A 100 000-year record of annual and seasonal rainfall and temperature for northwestern Australia based on a pollen record obtained offshore. Journal of Quaternary Science 21: 879-89.
,
A 100 000-year record of annual and seasonal rainfall and temperature for northwestern Australia based on a pollen record obtained offshore
,
Journal of Quaternary Science
, vol.
21
(pg.
879
-
89
)
Weckworth, B.V., Talbot, S.L. and Cook, J.A. 2010. Phylogeography of wolves (Canis lupus) in the Pacific Northwest. Journal of Mammalogy 91: 363-375.
,
Phylogeography of wolves (Canis lupus) in the Pacific Northwest
,
Journal of Mammalogy
, vol.
91
(pg.
363
-
375
)
Williams, M., Cook, E., Van der Kaars, S., Barrows, T., Shulmeister, J. and Kershaw, P. 2009. Glacial and deglacial climatic patterns in Australia and surrounding regions from 35 000 to 10 000 years ago reconstructed from terrestrial and near-shore proxy data. Quaternary Science Reviews 28: 2398-2419.
,
Glacial and deglacial climatic patterns in Australia and surrounding regions from 35 000 to 10 000 years ago reconstructed from terrestrial and near-shore proxy data
,
Quaternary Science Reviews
, vol.
28
(pg.
2398
-
2419
)
Willis, K. J. and Whittaker, R. J. 2000. Palaeoecology: The refugial debate. Science 287: 1406-1407.
,
Palaeoecology: The refugial debate
,
Science
, vol.
287
(pg.
1406
-
1407
)
Yokoyama, Y., Lambeck, K., De Deckker, P., Johnston, P. and Fifield, L. K. 2000. Timing of the Last Glacial Maximum from observed sea-level minima. Nature 406: 713-716.
,
Timing of the Last Glacial Maximum from observed sea-level minima
,
Nature
, vol.
406
(pg.
713
-
716
)
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