RICHARD Highton, an emeritus professor at the University of Maryland, is celebrated among herpetologists for his career of systematic inquiry into the salamander genus Plethodon (family Plethodontidae). He is also renowned for unusually complete geographic sampling. At the fourth Conference on the Biology of Plethodontid Salamanders in Highlands, North Carolina in 1998 (held in honor of Highton the year of his retirement), Steve Tilley gave a presentation on genetic diversity in Desmognathus salamanders. He started his talk with a slide of what the population-level sampling would look like if Highton had done it: like someone had blasted the range map with a shotgun. Tilley then presented his more modest sampling, and all the professors had a good laugh. Indeed, during the course of his career, Highton examined patterns of allozyme variation in >70,000 individuals from >1,500 populations throughout Plethodon. Moreover, he carried out numerous comprehensive studies of coloration, morphology,...

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