Research Zoologist and Curator of Amphibians and Reptiles, National Museum of Natural History, Smithsonian Institution

B.S. Biology, University of California at Los Angeles (Mentor George Gorman)

M.S. Zoology, San Diego State University (Advisor Richard Etheridge)

Ph.D. Zoology, University of California at Berkeley (Advisor David Wake)

Tilton Postdoctoral Fellow, California Academy of Sciences (Advisors Robert Drewes, Jacques Gauthier, and Alan Leviton)

Rufus Putnam Visiting Professor, Ohio University

Investigador Visitante, Instituto de Ecología (INECOL), Mexico

Edward P. Bass Distinguished Visiting Environmental Scholar, Yale University

My research involves the theory and practice of systematics. Most of my empirical research has been on iguanian lizards, including the true iguanas, the phrynosomatid sand lizards, hoplocercines, and anoles, and it most commonly involves phylogenetics, species delimitation, and studies of adaptive (esp. ecomorphological) evolution. My theoretical research has been devoted to developing a unified concept of species as well as a phylogenetic approach to biological nomenclature, and...

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